Higher education

• 1966-1969: B.Sc. Faculté des Sciences de Paris, Paris VII, France.
• 1971:  M.Sc. Institute of Molecular Biology, University Paris VII, France, under the supervision of Anne-Lise Haenni and Francois Chappeville.
• 1977: Teaching  Licensate in Chemistry, School of Education, Hebrew University of Jerusalem, Jerusalem, Israel
• 1978: Ph.D.  Department of Biological Chemistry, Hebrew University of Jerusalem, Israel, under the supervision of Nathan de Groot and Abraham Hochberg.
• 1977-80: Post-doctorate, New York State Institute for Basic Research in Mental Disabilities, Staten Island, New York, U.S.A.

Appointments

• 2016-todate: Professor Emeritus, Institute of plant Sciences and Genetics in Agriculture, Robert H. Smith Faculty of Agriculture, Food and Environment, The Hebrew University of Jerusalem, Israel.
• 1996-2016:   Institute of plant Sciences and Genetics in Agriculture, Robert H. Smith Faculty of Agriculture, Food and Environment, The Hebrew University of Jerusalem, Israel.
• 1989-96: Associate Professor, Department of Field and Vegetable Crops and Genetics, Faculty of Agriculture, The Hebrew University of Jerusalem, Rehovot, Israel.
• 1985-88: Senior Lecturer, Department of Field and Vegetable Crops and Genetics, Faculty of Agriculture, The Hebrew University of Jerusalem, Rehovot, Israel.
• 1980-84: Senior Research Scientist, Department of Cell Biology, The Weizmann Institute of Science, Rehovot, Israel.
• 1977-80: Research Scientist, New York State Institute for Basic Research in Mental Disabilities, Staten Island, New York, U.S.A.
• 1973-77: Assistant in Biochemistry, Department of Biological Chemistry, The Hebrew University of Jerusalem, Israel.
• 1971-72: Assistant in Biochemistry, Faculté de Médecine de Paris, France.